Factors for their
success - an analysis on their physiology
There are several factors responsibles
for the success of mosses and lichens in Alaskan region. The two groups
are remarkably similar in attributes beneficial in severe environments.
Both tend to occupy and create relatively favourable microenvironments.
Many species exhibit a broad response of net assimilation rate to temperature,
with maxima at 10-15oC but with positive net assimilation and
dark respiration continuing at or below 0oC. Light compensation
and saturation intensities are typically lower in mosses and lichens than
in vascular plants, and compensation levels decrease with temperature permitting
positive net assimilation under cool, low-light conditions.
Moreover, most species have little access
to soil moisture and lack an effective cuticle; this enables them to absorb
water through much of their surface but results in rapid water loss under
drying conditions. Yet, though the plants become inactive when dry, they
resume normal metabolism rapidly on remoistening. Thus, mosses and lichens
are thus adapted to switching rapidly between periods of metabolic activity
and rest, utilizing favourable conditions whenever they occur. This may
be facilitated by micromorphological features thought to facilitate simultaneous
uptake by moss leaves of both water and carbon dioxide. They also resist
frost very well by conferring tolerance of cytoplasmic dehydration resulting
from extracellullar ice formation.